| Holotype: Lepraria adhaerens K. Knudsen, Elix & Lendemer |
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Description: Thallus saxicolous, or muscicolous and lichenicolous over soil or rock, but not
usually growing directly on soil, diffuse, distinctly blue-gray, but sometimes with a dull brownish hue in
part, dispersed to contiguous, either patchy and thin (less than 0.5 mm thick), and spreading up to ca. 2 cm,
adhering to mosses, lichens or mineral grains of the substrate or on substrates where it is well-established
and dominant, forming a thick (less than 1 mm) areolate crust of fused granules with a loose upper layer,
divided by deep fissures between the areoles and covering areas up to several meters; always without lobes
or distinct rim; granules lumpy, adhering to one another, 40-100 µm diam., with a colorless outer layer
comprised of uneven and indistinct gelatinized hyphae or of irregular, paraplectenchymatous hyphae, ca. 2-
4(5) µm diam., one to two layers thick, surrounding an algal core; without projecting hyphae, but
frequently producing thin colorless ±thick-walled hyphae, 2-3 µm diam., acting as anchors or rhizines; the
chlorococcoid green algal, 10-14 µm in diam., often aggregating within the enlarged granules which
ultimately fragment into sticky agglomerations; without a medulla or lower surface, but older thalli forming
a lower necral layer of gelatinized granules which incorporates the remains of lichen or moss hosts mixed
with grains of the substrate; in several thalli non-lichenized algae were observed between the granules and
more rarely independent fungi or bacteria. Ascomata and conidiomata absent. Secondary metabolites: pannarin (major), zeorin (major or minor), sometimes atranorin (minor) |
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Type: USA: California: San Diego County: Torrey Pines State Park, 32° 54' 58"N, 117° 19' 56"W,
107 m, thin-soiled opening in maritime chaparral on sandstone bluffs, growing over Rinodina intermedia,
Lepraria xerophila, bryophytes and soil, 13.April.2005, K. Knudsen et al. 2700 (UCR!, holotype; ASU!, CANB!,
PH!, SD!, UGDA!, isotypes). |
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Image: Rolf Muertter. |
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