Lichen Holotypes
 
Holotype: Lepraria adhaerens K. Knudsen, Elix & Lendemer
  Description: Thallus saxicolous, or muscicolous and lichenicolous over soil or rock, but not usually growing directly on soil, diffuse, distinctly blue-gray, but sometimes with a dull brownish hue in part, dispersed to contiguous, either patchy and thin (less than 0.5 mm thick), and spreading up to ca. 2 cm, adhering to mosses, lichens or mineral grains of the substrate or on substrates where it is well-established and dominant, forming a thick (less than 1 mm) areolate crust of fused granules with a loose upper layer, divided by deep fissures between the areoles and covering areas up to several meters; always without lobes or distinct rim; granules lumpy, adhering to one another, 40-100 µm diam., with a colorless outer layer comprised of uneven and indistinct gelatinized hyphae or of irregular, paraplectenchymatous hyphae, ca. 2- 4(5) µm diam., one to two layers thick, surrounding an algal core; without projecting hyphae, but frequently producing thin colorless ±thick-walled hyphae, 2-3 µm diam., acting as anchors or rhizines; the chlorococcoid green algal, 10-14 µm in diam., often aggregating within the enlarged granules which ultimately fragment into sticky agglomerations; without a medulla or lower surface, but older thalli forming a lower necral layer of gelatinized granules which incorporates the remains of lichen or moss hosts mixed with grains of the substrate; in several thalli non-lichenized algae were observed between the granules and more rarely independent fungi or bacteria. Ascomata and conidiomata absent. Secondary metabolites: pannarin (major), zeorin (major or minor), sometimes atranorin (minor)
  Type: USA: California: San Diego County: Torrey Pines State Park, 32° 54' 58"N, 117° 19' 56"W,
107 m, thin-soiled opening in maritime chaparral on sandstone bluffs, growing over Rinodina intermedia,
Lepraria xerophila, bryophytes and soil, 13.April.2005, K. Knudsen et al. 2700 (UCR!, holotype; ASU!, CANB!,
PH!, SD!, UGDA!, isotypes).
 
  Image: Rolf Muertter.
 
   

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